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Formation of distinct structures called labiate processes found in some diatom species indicates an association with an organic complex called the labiate process apparatus. ÎҵķÒ룺¶ÀÌØ½á¹¹µÄÐγɣ¬¼´Ëùν´½Ðνø³Ì£¬ÔÚһЩ¹èÔåÎïÖÖÖб»·¢ÏÖ£¬Õâ±íÃ÷ÓëÓлú¸´ºÏÎïµÄ½áºÏ±»³ÆÎª´½Ðνø³ÌÆ÷¼þ¡£ ¹Ø¼üÊDz»ÖªµÀµÚ¶þ¸öcalledÊǸÉʲôÓõģ¬Ï£Íû¸ø¸ö½âÊÍ Circularization of the inner tube of the portula could involve protein-mediated molding of the SDV as could the positioning of the central and flanking pores. ÎҵķÒ룺¶ÔportulaÄڹܵĻ·»¯¿ÉÄÜÉæ¼°SDVµÄµ°°×½éµ¼³ÉÐÍ£¬¿É½«SDV¶¨Î»ÔÚÖÐÑëºÍÁ½²à¿×϶¡£ Ö÷ÒªÊDz»ÖªµÀÄǸö¡° as could¡±ÊǸÉʲôµÄ£¬ËùÒÔÕû¾ä»°¶¼Ðª²ËÁË£¬Ï£Íû¸ø¸ö½âÊÍ Õâ¾ä»°³ö×ÔChem. Rev. 2008, 108, 4855¨C4874 ÌâÄ¿ÊÇDiatoms, Biomineralization Processes, and Genomics ÄǶÎÎÄ×ÖÈçÏ£ºAnother example of mesoscale structure formation in T. pseudonana is formation of the portulae (Figure 14). The portulae appear to be designed for chitin extrusion123,124 through the outer opening. The inner and outer structures of the portulae are distinct (Figure 14a). The inner structure consists of a large central pore flanked on the sides by two horseshoe-shaped pores (Figure 14b). The outer pore is circular or oval shaped and buttressed by ribs extending from the valve surface (Figure 14c). The portulae are formed at a very early stage in valve synthesis, and the inner pore is created via circularization and extension of a cylindrical structure at an early stage in deposition (Figure 14d). The outer portion of the portula has an intermediate stage of formation where there is an overgrowth of silica surrounding the central inner pore and flanking subsidiary pores (Figure 14e). The completed structure consists of a tube of the inner pore and flanking subsidiary pores that open into a chamber on the outer valve face (Figure 14f). The chamber tapers to form the outer opening (Figure 14f). No details of portulae formation have been described in the literature, but these observations (Figure 14) suggest a highly controlled process involved in shaping the complicated structure. Formation of distinct structures called labiate processes found in some diatom species indicates an association with an organic complex called the labiate process apparatus.129 The labiate process apparatus has not been characterized at the molecular level, but a reasonable assumption is that it is a protein complex responsible for shaping of the SDV or silica. Perhaps a similar type of complex is involved in portulae formation. Circularization of the inner tube of the portula could involve protein-mediated molding of the SDV as could the positioning of the central and flanking pores. ·ÒëÈçÏ£º¹èÔåT. pseudonanaÖеÄÁíÒ»¸ö½é¹Û½á¹¹ÐγɵÄÀý×ÓÊÇportulaeµÄÐγɣ¨Í¼14£©¡£portulaeËÆºõÊÇΪ¼×¿ÇËØÍ¨¹ýÍⲿ³ö¿Ú¼·³ö¶øÉè¼ÆµÄ£¨123£¬124£©¡£portulaeµÄÄÚ²¿ºÍÍⲿ½á¹¹ÊDz»Í¬µÄ£¨Í¼14a£©¡£ÄÚ²¿½á¹¹°üÀ¨Ò»¸ö´óµÄÖÐÑë¿×϶£¬Á½²àÊÇÁ½¸öÂíÌãÐεĿף¨Í¼14b£©¡£Íâ²ã¿×ÊÇÔ²ÐλòÍÖÔ²Ðεģ¬ÓÉ´Ó¿ÇÃæ±íÃæ£¨Í¼14C£©ÑÓÉìµÄÀß¹ÇribÖ§³Å£¨Í¼14c£©¡£¸ÃportulaeÔÚ¿ÇÃæºÏ³ÉµÄÔçÆÚ¾ÍÐγÉÁË£¬¶øÄÚ²¿¿×϶ÊÇÔÚ³Á»ýÔçÆÚͨ¹ýÔ²ÖùÐνṹµÄ»·»¯ºÍÀ©Õ¹²úÉúµÄ£¨Í¼14d£©¡£¸ÃportulaµÄÍⲿ£¬ÓÐÒ»¸öÐγɵÄÖмä½×¶Î£¬Î§ÈÆÖÐÑëÄÚ²¿¿×϶ºÍÁ½²à¸½Êô¿×϶½øÐжþÑõ»¯¹èµÄÔöÉú£¨Í¼14e£©¡£ÒÑÍê³ÉµÄ½á¹¹ÓÉÁ½²¿·Ö¹¹³É£ºÄڿ׹ܡ¢ÏòÍâ¿ÇÃæÉϵÄÇ»ÊÒ³¨¿ªµÄÁ½²à¸½Êô¿×϶£¨Í¼14f£©¡£Ç»ÊÒÖð½¥±äϸÐγÉÍⲿ³ö¿Ú£¨Í¼14f£©¡£Ã»ÓÐportulaeÐγɵÄÏêÇéÔÚÎÄÏ×Öеõ½ÃèÊö£¬µ«ÕâЩ¹Û²ì×ÊÁÏ£¨Í¼14£©±íÃ÷ÁËÒ»¸öÓëÐγɸ´ÔӽṹÓйصĸ߶ȿØÖƹý³Ì¡£¶ÀÌØ½á¹¹µÄÐγɣ¬¼´Ëùν´½Ðνø³Ì£¬ÔÚһЩ¹èÔåÎïÖÖÖб»·¢ÏÖ£¬Õâ±íÃ÷ÓëÓлú¸´ºÏÎïµÄ½áºÏ±»³ÆÎª´½Ðνø³ÌÆ÷¼þ¡££¨129£©´½Ðνø³ÌÆ÷¼þÉÐδÔÚ·Ö×ÓˮƽÉϽøÐбíÕ÷£¬µ«ºÏÀíµÄ¼ÙÉèÊÇ£¬ËüÊÇÒ»µ°°×Öʸ´ºÏÎ¸ºÔðSDV»ò¶þÑõ»¯¹èµÄ³ÉÐΡ£Ò²ÐíÀàËÆµÄ¸´ºÏÎï»á²ÎÓëportulaeµÄÐγɡ£¶ÔportulaÄڹܵĻ·»¯¿ÉÄÜÉæ¼°SDVµÄµ°°×½éµ¼³ÉÐÍ£¬¿É½«SDV¶¨Î»ÔÚÖÐÑëºÍÁ½²à¿×϶¡£ |
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