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We developed a regression model for predicting branch needle mass based on branch size and whorl position within the crown (Mencuccini and Grace 1996)
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From https://en.wikipedia.org/wiki/Whorl
A whorl is a type of spiral or circular pattern.
Other meanings of whorl include:
Whorl (botany), used to describe the attachment of sepals, petals, leaves, or branches at a single point

or    https://en.wikipedia.org/wiki/Whorl_(botany)
In botany, a whorl is an arrangement of sepals, petals, leaves, stipules or branches that radiate from a single point and surround or wrap around the stem.[1][2] A whorl consists of at least three elements; a pair of opposite leaves is not called a whorl.
1. "whorl". thedictionary. Retrieved 19 August 2012.
2.  Lindley, John. A Glossary of Technical Terms Used in Botany, p.100, Bradbury and Evans, London, 1848.
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