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It is tempting to explore whether there are plant-specific advantages to storing C as organic acids rather than as carbohydrates when it is to be subsequently used for the assimilation or use of N. Interestingly, ammonium-preferring rice plant has a unique plant-type phosphoenolpyruvate carboxylase (PEPC), Osppc4, located in its chloroplasts that accounts for approximately one-third of total PEPC protein. Knockdown of Osppc4 suppresses ammonium assimilation and subsequent amino acid synthesis by decreasing organic acids, which are C-skeleton donors for these processes, suggesting that rice has a unique route for organic acid synthesis and that primary ammonium assimilation is not necessarily the same in all vascular plants

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